
Evolution and The History of Life in Deep Time
Course Description
Life on Earth carries a story written in molecules, bones, sediments, and branching lineages. This course guides you through that storyâfrom the origins of life and the rise of complex cells to multicellularity, animals, life on land, mass extinctions, and modern conservation. Youâll learn the mechanisms that power evolution (variation, heredity, selection), how molecular changes become anatomical diversity through development, and how scientists reconstruct the tree of life using evidence from DNA, fossils, and comparative morphology.
Weâll explore species concepts and speciation, phylogenetic methods (cladistics, parsimony, Bayesian inference), macroevolutionary dynamics (rates, clocks, adaptive radiations), and evoâdevo insights that reveal how new functions emerge from old parts. Youâll trace milestones such as abiogenesis, oxygenation, eukaryogenesis via endosymbiosis, multicellularity, and the Cambrian explosion, then follow plants, arthropods, and vertebrates as they adapt to life on landâsolving challenges like water loss, UV exposure, gravity, and airâbreathing.
Youâll examine extinction from background processes to the Big Five mass extinctions and the unfolding humanâdriven sixth, learning to separate ultimate drivers from proximal kill mechanisms and to recognize how recovery and resilience work over millions of years. In palaeoecology, youâll infer niches, gradients, communities, and depositional environments with care for taphonomic and sampling biases. In palaeobiogeography, youâll see how barriers and provinces shape distributions, how mixing events spur diversification, and how continents and oceans evolve through time (e.g., the Iapetus ocean). Finally, conservation palaeobiology connects deepâtime knowledge to present decisionsârevealing lag effects, thresholds, and multiâstressor interactions that shortâterm studies can miss, and underscoring the importance of protecting geoheritage.
By the end, evolution will be more than a topicâit will be your lens for understanding how life becomes, diversifies, collapses, and recovers, and how deepâtime context sharpens todayâs biodiversity and conservation challenges.
Course Features
Structured, short lectures with clear summaries for each topic.
Conceptâfirst explanations that make complex ideas approachable (e.g., homology vs. homoplasy; crownâ vs. stemâgroups).
Evidenceâdriven learning with case studies from fossils, DNA, ecology, and Earth history.
Critical thinking emphasis: uncertainty, support values, sampling and preservation biases, and methodological tradeâoffs.
Realâworld relevance through conservation palaeobiology and geoheritage.
FAQs
Is this course suitable for beginners? Yes. It begins with fundamentals and builds to advanced concepts in an accessible way.
Does it include both biology and geology? Yes. It integrates DNA, development, fossils, ecology, and Earth systems.
Will this help with teaching or communications? Absolutelyâclear definitions, modern frameworks, and narrative coherence are emphasized.
Is there a focus on current relevance? Yes. Conservation palaeobiology and geoheritage connect deepâtime insights to todayâs decisions.
Curriculum at a Glance
Section 1: Introduction â Foundations of evolution: variation, heredity, selection; genotypeâphenotype via development.
Section 2: Evolution â Theory vs. everyday language; consilience of evidence; scales of evolutionary change.
Section 3: Phylogeny â Taxonomy vs. systematics; cladistics; characters; parsimony and Bayesian inference; homology/homoplasy.
Section 4: Macroevolution â Adaptation and exaptation; clocks and rates; gradualism vs. punctuated equilibrium; adaptive radiations; evoâdevo.
Section 5: Milestones â Abiogenesis; LUCA; oxygenation; eukaryogenesis; multicellularity; animals and the Cambrian record.
Section 6: Life on Land â Precambrian terrestrial signals; plants, arthropods, vertebrates; forest ecosystems; MesozoicâCenozoic transitions.
Section 7: Extinction â Background vs. mass extinctions; ultimate/proximal causes; Big Five; the Sixth.
Section 8: Palaeoecology â Niches, gradients, community patterns; depositional environments; quantitative bias correction.
Section 9: Conservation Palaeobiology â Stressors and multiâstressor dynamics; lag effects; thresholds; geoheritage.
Section 10: Palaeobiogeography â Provinces, barriers, mixing events (BIMEs); vicariance; oceans and continents in motion.
Section 11: History of Palaeontology â From folklore and Enlightenment to modern molecular and computational approaches.
If youâre ready to understand how life evolves, diversifies, collapses, and recoversâand to use deepâtime insights to see todayâs biodiversity and conservation challenges more clearlyâenroll now and start reading Earthâs grand story with scientific confidence and curiosity.
Materials and videos provided under this course are on a Creative Commons (CC) License. Original work by Russel Gardwood has been used across the course.
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